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For most pens it doesn't really matter. Most pens depend on either capillary action or pressure and gravity has an insignificant effect.... as long as you use the pen enough. Leave a pen in disuse long enough and it will dry out no matter how you store it. This generally applies to fountain pens too. Though the time frames are shorter. Storing a fountain pen nib up is fine. Nib down may give you an extra day before it clogs up, but that's debatable. You should never store a fountain pen with shimmering inks nib down. Never. Also storing or carrying a fountain pen nib down increases the risk of leaks, especially if the pen suffers from creep.
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